Обсуждаем, страница 23 - гдз по биологии 11 класс учебник Пасечник, Каменский

Биология, 11 класс Учебник, авторы: Пасечник Владимир Васильевич, Каменский Андрей Александрович, Рубцов Александр Михайлович, Швецов Глеб Геннадьевич, Абовян Леван Арташесович, Гапонюк Зоя Георгиевна, издательство Просвещение, Москва, 2019

Авторы: Пасечник В. В., Каменский А. А., Рубцов А. М., Швецов Г. Г., Абовян Л. А., Гапонюк З. Г.

Тип: Учебник

Серия: линия жизни

Издательство: Просвещение

Год издания: 2019 - 2026

Уровень обучения: углублённый

ISBN: 978-5-09-087486-1 (2022)

Допущено Министерством просвещения Российской Федерации

Глава 1. Популяционно-видовой уровень. Параграф 3. Синтетическая теория эволюции. Проверьте себя - страница 23.

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                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:12 [
                            "id" => 5420
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Пасечник"
                            "field_bio" => null
                            "field_degree_rp" => "доктора педагогических наук"
                            "field_foreign_lang_name" => null
                            "field_foreign_lang_patronymic" => null
                            "field_foreign_lang_surname" => null
                            "field_name" => "Владимир"
                            "field_patronymic" => "Васильевич"
                            "field_surname_rp" => "Пасечника"
                          ]
                          #original: array:12 [
                            "id" => 5420
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Пасечник"
                            "field_bio" => null
                            "field_degree_rp" => "доктора педагогических наук"
                            "field_foreign_lang_name" => null
                            "field_foreign_lang_patronymic" => null
                            "field_foreign_lang_surname" => null
                            "field_name" => "Владимир"
                            "field_patronymic" => "Васильевич"
                            "field_surname_rp" => "Пасечника"
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                        1 => App\Models\Term {#1140
                          #connection: "mysql"
                          #table: "terms"
                          #primaryKey: "id"
                          #keyType: "int"
                          +incrementing: false
                          #with: []
                          #withCount: []
                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:12 [
                            "id" => 4539
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Каменский"
                            "field_bio" => null
                            "field_degree_rp" => "доктора биологических наук"
                            "field_foreign_lang_name" => null
                            "field_foreign_lang_patronymic" => null
                            "field_foreign_lang_surname" => null
                            "field_name" => "Андрей"
                            "field_patronymic" => "Александрович"
                            "field_surname_rp" => "Каменского"
                          ]
                          #original: array:12 [
                            "id" => 4539
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Каменский"
                            "field_bio" => null
                            "field_degree_rp" => "доктора биологических наук"
                            "field_foreign_lang_name" => null
                            "field_foreign_lang_patronymic" => null
                            "field_foreign_lang_surname" => null
                            "field_name" => "Андрей"
                            "field_patronymic" => "Александрович"
                            "field_surname_rp" => "Каменского"
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                        2 => App\Models\Term {#1141
                          #connection: "mysql"
                          #table: "terms"
                          #primaryKey: "id"
                          #keyType: "int"
                          +incrementing: false
                          #with: []
                          #withCount: []
                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:12 [
                            "id" => 5627
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Рубцов"
                            "field_bio" => null
                            "field_degree_rp" => "доктора биологических наук"
                            "field_foreign_lang_name" => null
                            "field_foreign_lang_patronymic" => null
                            "field_foreign_lang_surname" => null
                            "field_name" => "Александр"
                            "field_patronymic" => "Михайлович"
                            "field_surname_rp" => "Рубцова"
                          ]
                          #original: array:12 [
                            "id" => 5627
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Рубцов"
                            "field_bio" => null
                            "field_degree_rp" => "доктора биологических наук"
                            "field_foreign_lang_name" => null
                            "field_foreign_lang_patronymic" => null
                            "field_foreign_lang_surname" => null
                            "field_name" => "Александр"
                            "field_patronymic" => "Михайлович"
                            "field_surname_rp" => "Рубцова"
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                        3 => App\Models\Term {#1142
                          #connection: "mysql"
                          #table: "terms"
                          #primaryKey: "id"
                          #keyType: "int"
                          +incrementing: false
                          #with: []
                          #withCount: []
                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:12 [
                            "id" => 6304
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Швецов"
                            "field_bio" => null
                            "field_degree_rp" => "кандидата педагогических наук"
                            "field_foreign_lang_name" => null
                            "field_foreign_lang_patronymic" => null
                            "field_foreign_lang_surname" => null
                            "field_name" => "Глеб"
                            "field_patronymic" => "Геннадьевич"
                            "field_surname_rp" => null
                          ]
                          #original: array:12 [
                            "id" => 6304
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Швецов"
                            "field_bio" => null
                            "field_degree_rp" => "кандидата педагогических наук"
                            "field_foreign_lang_name" => null
                            "field_foreign_lang_patronymic" => null
                            "field_foreign_lang_surname" => null
                            "field_name" => "Глеб"
                            "field_patronymic" => "Геннадьевич"
                            "field_surname_rp" => null
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                        4 => App\Models\Term {#1143
                          #connection: "mysql"
                          #table: "terms"
                          #primaryKey: "id"
                          #keyType: "int"
                          +incrementing: false
                          #with: []
                          #withCount: []
                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:12 [
                            "id" => 6898
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Абовян"
                            "field_bio" => null
                            "field_degree_rp" => null
                            "field_foreign_lang_name" => null
                            "field_foreign_lang_patronymic" => null
                            "field_foreign_lang_surname" => null
                            "field_name" => "Леван"
                            "field_patronymic" => "Арташесович"
                            "field_surname_rp" => null
                          ]
                          #original: array:12 [
                            "id" => 6898
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Абовян"
                            "field_bio" => null
                            "field_degree_rp" => null
                            "field_foreign_lang_name" => null
                            "field_foreign_lang_patronymic" => null
                            "field_foreign_lang_surname" => null
                            "field_name" => "Леван"
                            "field_patronymic" => "Арташесович"
                            "field_surname_rp" => null
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                        5 => App\Models\Term {#1144
                          #connection: "mysql"
                          #table: "terms"
                          #primaryKey: "id"
                          #keyType: "int"
                          +incrementing: false
                          #with: []
                          #withCount: []
                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:12 [
                            "id" => 4086
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Гапонюк"
                            "field_bio" => null
                            "field_degree_rp" => "кандидата педагогических наук"
                            "field_foreign_lang_name" => null
                            "field_foreign_lang_patronymic" => null
                            "field_foreign_lang_surname" => null
                            "field_name" => "Зоя"
                            "field_patronymic" => "Георгиевна"
                            "field_surname_rp" => null
                          ]
                          #original: array:12 [
                            "id" => 4086
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Гапонюк"
                            "field_bio" => null
                            "field_degree_rp" => "кандидата педагогических наук"
                            "field_foreign_lang_name" => null
                            "field_foreign_lang_patronymic" => null
                            "field_foreign_lang_surname" => null
                            "field_name" => "Зоя"
                            "field_patronymic" => "Георгиевна"
                            "field_surname_rp" => null
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                      ]
                      #escapeWhenCastingToString: false
                    }
                    "field_author_foreign" => Illuminate\Database\Eloquent\Collection {#1145
                      #items: []
                      #escapeWhenCastingToString: false
                    }
                    "field_book_type" => Illuminate\Database\Eloquent\Collection {#1146
                      #items: array:1 [
                        0 => App\Models\Term {#1147
                          #connection: "mysql"
                          #table: "terms"
                          #primaryKey: "id"
                          #keyType: "int"
                          +incrementing: false
                          #with: []
                          #withCount: []
                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:10 [
                            "id" => 6671
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Учебник"
                            "field_book_type_foreign" => null
                            "field_cases" => array:6 [ …6]
                            "field_plural_form" => null
                            "field_short_name" => null
                            "field_short_name_foreign" => null
                            "field_translit" => "uchebnik"
                          ]
                          #original: array:10 [
                            "id" => 6671
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Учебник"
                            "field_book_type_foreign" => null
                            "field_cases" => array:6 [ …6]
                            "field_plural_form" => null
                            "field_short_name" => null
                            "field_short_name_foreign" => null
                            "field_translit" => "uchebnik"
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                      ]
                      #escapeWhenCastingToString: false
                    }
                    "field_country" => Illuminate\Database\Eloquent\Collection {#1148
                      #items: array:1 [
                        0 => App\Models\Term {#1149
                          #connection: "mysql"
                          #table: "terms"
                          #primaryKey: "id"
                          #keyType: "int"
                          +incrementing: false
                          #with: []
                          #withCount: []
                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:6 [
                            "id" => 9
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Россия"
                            "field_cases" => array:6 [ …6]
                            "field_translit" => "rossiya"
                          ]
                          #original: array:6 [
                            "id" => 9
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Россия"
                            "field_cases" => array:6 [ …6]
                            "field_translit" => "rossiya"
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                      ]
                      #escapeWhenCastingToString: false
                    }
                    "field_city" => Illuminate\Database\Eloquent\Collection {#1150
                      #items: array:1 [
                        0 => App\Models\Term {#1151
                          #connection: "mysql"
                          #table: "terms"
                          #primaryKey: "id"
                          #keyType: "int"
                          +incrementing: false
                          #with: []
                          #withCount: []
                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:6 [
                            "id" => 15
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Москва"
                            "field_cases" => array:6 [ …6]
                            "field_translit" => "moskva"
                          ]
                          #original: array:6 [
                            "id" => 15
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "Москва"
                            "field_cases" => array:6 [ …6]
                            "field_translit" => "moskva"
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                      ]
                      #escapeWhenCastingToString: false
                    }
                    "field_series" => Illuminate\Database\Eloquent\Collection {#1152
                      #items: array:1 [
                        0 => App\Models\Term {#1153
                          #connection: "mysql"
                          #table: "terms"
                          #primaryKey: "id"
                          #keyType: "int"
                          +incrementing: false
                          #with: []
                          #withCount: []
                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:8 [
                            "id" => 4725
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "линия жизни"
                            "field_cases" => null
                            "field_foreign_lang_series" => null
                            "field_transcription_eng" => null
                            "field_translit" => "liniya zhizni"
                          ]
                          #original: array:8 [
                            "id" => 4725
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "линия жизни"
                            "field_cases" => null
                            "field_foreign_lang_series" => null
                            "field_transcription_eng" => null
                            "field_translit" => "liniya zhizni"
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                      ]
                      #escapeWhenCastingToString: false
                    }
                    "field_umk" => Illuminate\Database\Eloquent\Collection {#1154
                      #items: []
                      #escapeWhenCastingToString: false
                    }
                    "field_level_of_education" => Illuminate\Database\Eloquent\Collection {#1155
                      #items: array:1 [
                        0 => App\Models\Term {#1156
                          #connection: "mysql"
                          #table: "terms"
                          #primaryKey: "id"
                          #keyType: "int"
                          +incrementing: false
                          #with: []
                          #withCount: []
                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:6 [
                            "id" => 39
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "углублённый"
                            "field_cases" => array:6 [ …6]
                            "field_translit" => "uglublennyy"
                          ]
                          #original: array:6 [
                            "id" => 39
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "углублённый"
                            "field_cases" => array:6 [ …6]
                            "field_translit" => "uglublennyy"
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                      ]
                      #escapeWhenCastingToString: false
                    }
                    "field_standart_of_education" => Illuminate\Database\Eloquent\Collection {#1157
                      #items: array:1 [
                        0 => App\Models\Term {#1158
                          #connection: "mysql"
                          #table: "terms"
                          #primaryKey: "id"
                          #keyType: "int"
                          +incrementing: false
                          #with: []
                          #withCount: []
                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:10 [
                            "id" => 6706
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "ФГОС (старый)"
                            "field_cases" => array:6 [ …6]
                            "field_color" => null
                            "field_end_year" => "2022"
                            "field_start_year" => "2019"
                            "field_translit" => "fgos-old"
                            "field_type" => null
                          ]
                          #original: array:10 [
                            "id" => 6706
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "ФГОС (старый)"
                            "field_cases" => array:6 [ …6]
                            "field_color" => null
                            "field_end_year" => "2022"
                            "field_start_year" => "2019"
                            "field_translit" => "fgos-old"
                            "field_type" => null
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                      ]
                      #escapeWhenCastingToString: false
                    }
                    "field_publication_number" => "4"
                    "field_publication_type" => Illuminate\Database\Eloquent\Collection {#1159
                      #items: array:1 [
                        0 => App\Models\Term {#1160
                          #connection: "mysql"
                          #table: "terms"
                          #primaryKey: "id"
                          #keyType: "int"
                          +incrementing: false
                          #with: []
                          #withCount: []
                          +preventsLazyLoading: false
                          #perPage: 15
                          +exists: true
                          +wasRecentlyCreated: false
                          #escapeWhenCastingToString: false
                          #attributes: array:6 [
                            "id" => 33
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "стереотипное"
                            "field_cases" => array:6 [ …6]
                            "field_translit" => "stereotipnoe"
                          ]
                          #original: array:6 [
                            "id" => 33
                            "created_at" => "2026-04-10 13:58:26"
                            "updated_at" => null
                            "name" => "стереотипное"
                            "field_cases" => array:6 [ …6]
                            "field_translit" => "stereotipnoe"
                          ]
                          #changes: []
                          #casts: []
                          #classCastCache: []
                          #attributeCastCache: []
                          #dateFormat: null
                          #appends: []
                          #dispatchesEvents: []
                          #observables: []
                          #relations: []
                          #touches: []
                          +timestamps: true
                          +usesUniqueIds: false
                          #hidden: []
                          #visible: []
                          #fillable: []
                          #guarded: array:1 [
                            0 => "*"
                          ]
                        }
                      ]
                      #escapeWhenCastingToString: false
                    }
                    "field_under_the_edition" => Illuminate\Database\Eloquent\Collection {#1161
                      #items: array:1 [
                        0 => App\Models\Term {#1139}
                      ]
                      #escapeWhenCastingToString: false
                    }
                    "field_under_the_edition_degree" => "доктора педагогических наук"
                    "field_cover_description" => null
                    "field_publication_year" => "2019"
                    "field_publication_year_until" => null
                    "field_part" => ""
                    "field_part_writing" => ""
                    "field_second_foreign_language" => null
                    "field_for_whom" => "Учебник для общеобразовательных организаций"
                    "field_allowed" => "Допущено Министерством просвещения Российской Федерации"
                    "field_reserve_field" => null
                    "field_link_to_source" => null
                    "field_tasks_count" => "387"
                    "field_priority" => null
                    "field_default_folder" => "/biologija_11/pasechnik-u19/"
                    "field_isbn" => "978-5-09-087486-1 (2022)"
                    "field_cover" => array:1 [
                      0 => "/media/biologija_11/pasechnik-u19/covers/cover1.webp?ts=1759691562"
                    ]
                    "field_cover_alts" => array:1 [
                      0 => ""
                    ]
                    "field_covers" => array:1 [
                      0 => array:4 [
                        "path" => "/media/biologija_11/pasechnik-u19/covers/cover1.webp?ts=1759691562"
                        "alt" => ""
                        "width" => "1200"
                        "height" => "1584"
                      ]
                    ]
                    "field_popular_book" => "0"
                    "field_recommended_books" => Illuminate\Database\Eloquent\Collection {#1162
                      #items: []
                      #escapeWhenCastingToString: false
                    }
                    "field_new_book" => Illuminate\Database\Eloquent\Collection {#1163
                      #items: []
                      #escapeWhenCastingToString: false
                    }
                    "field_old_book" => Illuminate\Database\Eloquent\Collection {#1164
                      #items: []
                      #escapeWhenCastingToString: false
                    }
                    "field_url" => "/11-klass/biologija/pasechnik"
                    "field_cover_color" => Illuminate\Database\Eloquent\Collection {#1165
                      #items: []
                      #escapeWhenCastingToString: false
                    }
                    "field_metatags_title" => null
                    "field_metatags_description" => null
                    "field_h1" => null
                    "field_description_top" => null
                    "field_description_bottom" => null
                    "breadcrumbs" => array:3 [
                      "class" => 382
                      "subject" => 552
                      "class_subject" => 462
                    ]
                  ]
                  #original: array:50 [
                    "id" => 2964
                    "created_at" => "2026-04-10 13:58:26"
                    "updated_at" => null
                    "field_tree_default_status" => "tasks"
                    "field_pages_status" => null
                    "field_subject" => Illuminate\Database\Eloquent\Collection {#1133}
                    "field_class" => Illuminate\Database\Eloquent\Collection {#1134}
                    "field_publisher" => Illuminate\Database\Eloquent\Collection {#1136}
                    "field_author" => Illuminate\Database\Eloquent\Collection {#1138}
                    "field_author_foreign" => Illuminate\Database\Eloquent\Collection {#1145}
                    "field_book_type" => Illuminate\Database\Eloquent\Collection {#1146}
                    "field_country" => Illuminate\Database\Eloquent\Collection {#1148}
                    "field_city" => Illuminate\Database\Eloquent\Collection {#1150}
                    "field_series" => Illuminate\Database\Eloquent\Collection {#1152}
                    "field_umk" => Illuminate\Database\Eloquent\Collection {#1154}
                    "field_level_of_education" => Illuminate\Database\Eloquent\Collection {#1155}
                    "field_standart_of_education" => Illuminate\Database\Eloquent\Collection {#1157}
                    "field_publication_number" => "4"
                    "field_publication_type" => Illuminate\Database\Eloquent\Collection {#1159}
                    "field_under_the_edition" => Illuminate\Database\Eloquent\Collection {#1161}
                    "field_under_the_edition_degree" => "доктора педагогических наук"
                    "field_cover_description" => null
                    "field_publication_year" => "2019"
                    "field_publication_year_until" => null
                    "field_part" => ""
                    "field_part_writing" => ""
                    "field_second_foreign_language" => null
                    "field_for_whom" => "Учебник для общеобразовательных организаций"
                    "field_allowed" => "Допущено Министерством просвещения Российской Федерации"
                    "field_reserve_field" => null
                    "field_link_to_source" => null
                    "field_tasks_count" => "387"
                    "field_priority" => null
                    "field_default_folder" => "/biologija_11/pasechnik-u19/"
                    "field_isbn" => "978-5-09-087486-1 (2022)"
                    "field_cover" => array:1 [
                      0 => "/media/biologija_11/pasechnik-u19/covers/cover1.webp?ts=1759691562"
                    ]
                    "field_cover_alts" => array:1 [
                      0 => ""
                    ]
                    "field_covers" => array:1 [
                      0 => array:4 [
                        "path" => "/media/biologija_11/pasechnik-u19/covers/cover1.webp?ts=1759691562"
                        "alt" => ""
                        "width" => "1200"
                        "height" => "1584"
                      ]
                    ]
                    "field_popular_book" => "0"
                    "field_recommended_books" => Illuminate\Database\Eloquent\Collection {#1162}
                    "field_new_book" => Illuminate\Database\Eloquent\Collection {#1163}
                    "field_old_book" => Illuminate\Database\Eloquent\Collection {#1164}
                    "field_url" => "/11-klass/biologija/pasechnik"
                    "field_cover_color" => Illuminate\Database\Eloquent\Collection {#1165}
                    "field_metatags_title" => null
                    "field_metatags_description" => null
                    "field_h1" => null
                    "field_description_top" => null
                    "field_description_bottom" => null
                    "breadcrumbs" => array:3 [
                      "class" => 382
                      "subject" => 552
                      "class_subject" => 462
                    ]
                  ]
                  #changes: []
                  #casts: []
                  #classCastCache: []
                  #attributeCastCache: []
                  #dateFormat: null
                  #appends: []
                  #dispatchesEvents: []
                  #observables: []
                  #relations: []
                  #touches: []
                  +timestamps: true
                  +usesUniqueIds: false
                  #hidden: []
                  #visible: []
                  #fillable: []
                  #guarded: array:1 [
                    0 => "*"
                  ]
                }
              ]
              #escapeWhenCastingToString: false
            }
            "branch_parent" => Illuminate\Database\Eloquent\Collection {#1166
              #items: []
              #escapeWhenCastingToString: false
            }
          ]
          #original: array:24 [
            "id" => 1450905
            "created_at" => "2026-04-10 13:58:26"
            "updated_at" => null
            "field_display_title" => "Популяционно-видовой уровень"
            "field_branch_order" => "1"
            "field_url" => null
            "field_branch_type" => Illuminate\Database\Eloquent\Collection {#1132}
            "field_page_start" => "5"
            "field_branch_display" => "0"
            "field_branch_expanded" => "0"
            "field_display_branch_in_title" => "1"
            "field_display_task_interval" => "0"
            "field_display_branch_page" => "1"
            "field_branch_title_in_content" => "0"
            "field_navigation_title" => null
            "field_metatags_title" => null
            "field_metatags_description" => null
            "field_h1" => null
            "field_description_top" => null
            "field_description_bottom" => null
            "field_branch_cover" => []
            "field_branch_covers" => []
            "book" => Illuminate\Database\Eloquent\Collection {#1168}
            "branch_parent" => Illuminate\Database\Eloquent\Collection {#1166}
          ]
          #changes: []
          #casts: []
          #classCastCache: []
          #attributeCastCache: []
          #dateFormat: null
          #appends: []
          #dispatchesEvents: []
          #observables: []
          #relations: []
          #touches: []
          +timestamps: true
          +usesUniqueIds: false
          #hidden: []
          #visible: []
          #fillable: []
          #guarded: array:1 [
            0 => "*"
          ]
        }
        1 => App\Models\Branch {#1167
          #connection: "mysql"
          #table: "branches"
          #primaryKey: "id"
          #keyType: "int"
          +incrementing: true
          #with: []
          #withCount: []
          +preventsLazyLoading: false
          #perPage: 15
          +exists: true
          +wasRecentlyCreated: false
          #escapeWhenCastingToString: false
          #attributes: array:24 [
            "id" => 1450912
            "created_at" => "2026-04-10 13:58:26"
            "updated_at" => null
            "field_display_title" => "Синтетическая теория эволюции"
            "field_branch_order" => "3"
            "field_url" => null
            "field_branch_type" => Illuminate\Database\Eloquent\Collection {#1172
              #items: array:1 [
                0 => App\Models\Term {#1169
                  #connection: "mysql"
                  #table: "terms"
                  #primaryKey: "id"
                  #keyType: "int"
                  +incrementing: false
                  #with: []
                  #withCount: []
                  +preventsLazyLoading: false
                  #perPage: 15
                  +exists: true
                  +wasRecentlyCreated: false
                  #escapeWhenCastingToString: false
                  #attributes: array:7 [
                    "id" => 32
                    "created_at" => "2026-04-10 13:58:26"
                    "updated_at" => null
                    "name" => "Параграф"
                    "field_cases" => array:6 [ …6]
                    "field_page_check_not_needed" => null
                    "field_short_name" => "§"
                  ]
                  #original: array:7 [
                    "id" => 32
                    "created_at" => "2026-04-10 13:58:26"
                    "updated_at" => null
                    "name" => "Параграф"
                    "field_cases" => array:6 [ …6]
                    "field_page_check_not_needed" => null
                    "field_short_name" => "§"
                  ]
                  #changes: []
                  #casts: []
                  #classCastCache: []
                  #attributeCastCache: []
                  #dateFormat: null
                  #appends: []
                  #dispatchesEvents: []
                  #observables: []
                  #relations: []
                  #touches: []
                  +timestamps: true
                  +usesUniqueIds: false
                  #hidden: []
                  #visible: []
                  #fillable: []
                  #guarded: array:1 [
                    0 => "*"
                  ]
                }
              ]
              #escapeWhenCastingToString: false
            }
            "field_page_start" => "20"
            "field_branch_display" => "0"
            "field_branch_expanded" => "0"
            "field_display_branch_in_title" => "1"
            "field_display_task_interval" => "0"
            "field_display_branch_page" => "1"
            "field_branch_title_in_content" => "0"
            "field_navigation_title" => null
            "field_metatags_title" => null
            "field_metatags_description" => null
            "field_h1" => null
            "field_description_top" => null
            "field_description_bottom" => null
            "field_branch_cover" => []
            "field_branch_covers" => []
            "book" => Illuminate\Database\Eloquent\Collection {#1171
              #items: array:1 [
                0 => App\Models\Book {#1131}
              ]
              #escapeWhenCastingToString: false
            }
            "branch_parent" => Illuminate\Database\Eloquent\Collection {#1170
              #items: array:1 [
                0 => App\Models\Branch {#1035}
              ]
              #escapeWhenCastingToString: false
            }
          ]
          #original: array:24 [
            "id" => 1450912
            "created_at" => "2026-04-10 13:58:26"
            "updated_at" => null
            "field_display_title" => "Синтетическая теория эволюции"
            "field_branch_order" => "3"
            "field_url" => null
            "field_branch_type" => Illuminate\Database\Eloquent\Collection {#1172}
            "field_page_start" => "20"
            "field_branch_display" => "0"
            "field_branch_expanded" => "0"
            "field_display_branch_in_title" => "1"
            "field_display_task_interval" => "0"
            "field_display_branch_page" => "1"
            "field_branch_title_in_content" => "0"
            "field_navigation_title" => null
            "field_metatags_title" => null
            "field_metatags_description" => null
            "field_h1" => null
            "field_description_top" => null
            "field_description_bottom" => null
            "field_branch_cover" => []
            "field_branch_covers" => []
            "book" => Illuminate\Database\Eloquent\Collection {#1171}
            "branch_parent" => Illuminate\Database\Eloquent\Collection {#1170}
          ]
          #changes: []
          #casts: []
          #classCastCache: []
          #attributeCastCache: []
          #dateFormat: null
          #appends: []
          #dispatchesEvents: []
          #observables: []
          #relations: []
          #touches: []
          +timestamps: true
          +usesUniqueIds: false
          #hidden: []
          #visible: []
          #fillable: []
          #guarded: array:1 [
            0 => "*"
          ]
        }
        2 => App\Models\Branch {#1173
          #connection: "mysql"
          #table: "branches"
          #primaryKey: "id"
          #keyType: "int"
          +incrementing: true
          #with: []
          #withCount: []
          +preventsLazyLoading: false
          #perPage: 15
          +exists: true
          +wasRecentlyCreated: false
          #escapeWhenCastingToString: false
          #attributes: array:24 [
            "id" => 1450914
            "created_at" => "2026-04-10 13:58:26"
            "updated_at" => null
            "field_display_title" => "Проверьте себя"
            "field_branch_order" => null
            "field_url" => null
            "field_branch_type" => Illuminate\Database\Eloquent\Collection {#1174
              #items: []
              #escapeWhenCastingToString: false
            }
            "field_page_start" => "23"
            "field_branch_display" => "0"
            "field_branch_expanded" => "0"
            "field_display_branch_in_title" => "1"
            "field_display_task_interval" => "0"
            "field_display_branch_page" => "1"
            "field_branch_title_in_content" => "0"
            "field_navigation_title" => null
            "field_metatags_title" => null
            "field_metatags_description" => null
            "field_h1" => null
            "field_description_top" => null
            "field_description_bottom" => null
            "field_branch_cover" => []
            "field_branch_covers" => []
            "book" => Illuminate\Database\Eloquent\Collection {#1175
              #items: array:1 [
                0 => App\Models\Book {#1131}
              ]
              #escapeWhenCastingToString: false
            }
            "branch_parent" => Illuminate\Database\Eloquent\Collection {#1184
              #items: array:1 [
                0 => App\Models\Branch {#1183
                  #connection: "mysql"
                  #table: "branches"
                  #primaryKey: "id"
                  #keyType: "int"
                  +incrementing: true
                  #with: []
                  #withCount: []
                  +preventsLazyLoading: false
                  #perPage: 15
                  +exists: true
                  +wasRecentlyCreated: false
                  #escapeWhenCastingToString: false
                  #attributes: array:24 [
                    "id" => 1450912
                    "created_at" => "2026-04-10 13:58:26"
                    "updated_at" => null
                    "field_display_title" => "Синтетическая теория эволюции"
                    "field_branch_order" => "3"
                    "field_url" => null
                    "field_branch_type" => Illuminate\Database\Eloquent\Collection {#1185 …2}
                    "field_page_start" => "20"
                    "field_branch_display" => "0"
                    "field_branch_expanded" => "0"
                    "field_display_branch_in_title" => "1"
                    "field_display_task_interval" => "0"
                    "field_display_branch_page" => "1"
                    "field_branch_title_in_content" => "0"
                    "field_navigation_title" => null
                    "field_metatags_title" => null
                    "field_metatags_description" => null
                    "field_h1" => null
                    "field_description_top" => null
                    "field_description_bottom" => null
                    "field_branch_cover" => []
                    "field_branch_covers" => []
                    "book" => Illuminate\Database\Eloquent\Collection {#1186 …2}
                    "branch_parent" => Illuminate\Database\Eloquent\Collection {#1187 …2}
                  ]
                  #original: array:24 [
                    "id" => 1450912
                    "created_at" => "2026-04-10 13:58:26"
                    "updated_at" => null
                    "field_display_title" => "Синтетическая теория эволюции"
                    "field_branch_order" => "3"
                    "field_url" => null
                    "field_branch_type" => Illuminate\Database\Eloquent\Collection {#1185 …2}
                    "field_page_start" => "20"
                    "field_branch_display" => "0"
                    "field_branch_expanded" => "0"
                    "field_display_branch_in_title" => "1"
                    "field_display_task_interval" => "0"
                    "field_display_branch_page" => "1"
                    "field_branch_title_in_content" => "0"
                    "field_navigation_title" => null
                    "field_metatags_title" => null
                    "field_metatags_description" => null
                    "field_h1" => null
                    "field_description_top" => null
                    "field_description_bottom" => null
                    "field_branch_cover" => []
                    "field_branch_covers" => []
                    "book" => Illuminate\Database\Eloquent\Collection {#1186 …2}
                    "branch_parent" => Illuminate\Database\Eloquent\Collection {#1187 …2}
                  ]
                  #changes: []
                  #casts: []
                  #classCastCache: []
                  #attributeCastCache: []
                  #dateFormat: null
                  #appends: []
                  #dispatchesEvents: []
                  #observables: []
                  #relations: []
                  #touches: []
                  +timestamps: true
                  +usesUniqueIds: false
                  #hidden: []
                  #visible: []
                  #fillable: []
                  #guarded: array:1 [
                    0 => "*"
                  ]
                }
              ]
              #escapeWhenCastingToString: false
            }
          ]
          #original: array:24 [
            "id" => 1450914
            "created_at" => "2026-04-10 13:58:26"
            "updated_at" => null
            "field_display_title" => "Проверьте себя"
            "field_branch_order" => null
            "field_url" => null
            "field_branch_type" => Illuminate\Database\Eloquent\Collection {#1174}
            "field_page_start" => "23"
            "field_branch_display" => "0"
            "field_branch_expanded" => "0"
            "field_display_branch_in_title" => "1"
            "field_display_task_interval" => "0"
            "field_display_branch_page" => "1"
            "field_branch_title_in_content" => "0"
            "field_navigation_title" => null
            "field_metatags_title" => null
            "field_metatags_description" => null
            "field_h1" => null
            "field_description_top" => null
            "field_description_bottom" => null
            "field_branch_cover" => []
            "field_branch_covers" => []
            "book" => Illuminate\Database\Eloquent\Collection {#1175}
            "branch_parent" => Illuminate\Database\Eloquent\Collection {#1184}
          ]
          #changes: []
          #casts: []
          #classCastCache: []
          #attributeCastCache: []
          #dateFormat: null
          #appends: []
          #dispatchesEvents: []
          #observables: []
          #relations: []
          #touches: []
          +timestamps: true
          +usesUniqueIds: false
          #hidden: []
          #visible: []
          #fillable: []
          #guarded: array:1 [
            0 => "*"
          ]
        }
      ]
      #escapeWhenCastingToString: false
    }
    "content" => Illuminate\Database\Eloquent\Collection {#1111
      #items: array:3 [
        0 => App\Models\Element {#1103
          #connection: "mysql"
          #table: "elements"
          #primaryKey: "id"
          #keyType: "int"
          +incrementing: true
          #with: []
          #withCount: []
          +preventsLazyLoading: false
          #perPage: 15
          +exists: true
          +wasRecentlyCreated: false
          #escapeWhenCastingToString: false
          #attributes: array:7 [
            "id" => 1741151
            "created_at" => "2026-04-10 13:58:26"
            "updated_at" => null
            "edition" => Illuminate\Database\Eloquent\Collection {#1094
              #items: array:1 [
                0 => App\Models\Edition {#1100
                  #connection: "mysql"
                  #table: "editions"
                  #primaryKey: "id"
                  #keyType: "int"
                  +incrementing: false
                  #with: []
                  #withCount: []
                  +preventsLazyLoading: false
                  #perPage: 15
                  +exists: true
                  +wasRecentlyCreated: false
                  #escapeWhenCastingToString: false
                  #attributes: array:21 [
                    "id" => 6254
                    "created_at" => "2026-04-10 13:58:26"
                    "updated_at" => null
                    "title" => "Условие"
                    "field_order" => "1"
                    "field_publisher" => Illuminate\Database\Eloquent\Collection {#1098 …2}
                    "field_content_type" => "free"
                    "field_content_mode" => "text, image"
                    "field_page_content_mode" => "image"
                    "field_content_text_checked" => "0"
                    "field_page_content_text_checked" => "0"
                    "field_solution_author" => "Автор"
                    "field_moderator" => "pasha"
                    "field_edition_type" => "statement"
                    "field_root_dir" => "0-"
                    "field_responsible" => Illuminate\Database\Eloquent\Collection {#1099 …2}
                    "field_comment" => null
                    "field_similar_book" => Illuminate\Database\Eloquent\Collection {#1096 …2}
                    "field_process_formula" => ""
                    "field_edition_group" => Illuminate\Database\Eloquent\Collection {#1097 …2}
                    "field_content_source" => null
                  ]
                  #original: array:21 [
                    "id" => 6254
                    "created_at" => "2026-04-10 13:58:26"
                    "updated_at" => null
                    "title" => "Условие"
                    "field_order" => "1"
                    "field_publisher" => Illuminate\Database\Eloquent\Collection {#1098 …2}
                    "field_content_type" => "free"
                    "field_content_mode" => "text, image"
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            "text" => "<p>Прочитайте статью об эксперименте группы Ленски в рубрике «Это интересно», после чего обсудите с одноклассниками и с преподавателем следующие вопросы:</p><p><strong>1.</strong> Можно ли считать, что эксперимент группы Ленски действительно привёл весомые доказательства в пользу справедливости синтетической теории эволюции, или всё-таки результатов этого эксперимента недостаточно для того, чтобы считать эти положения доказанными? Приведите аргументы в пользу обеих точек зрения.</p><p><strong>2.</strong> Противники эволюционной теории достаточно часто говорят о том, что новый признак у организма не может появиться в результате мутации. Как вы думаете, можно ли опровергнуть данное утверждение, приведя в пример некоторые результаты эксперимента группы Ленски?</p>"
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            "text" => "<p>Прочитайте статью об эксперименте группы Ленски в рубрике «Это интересно», после чего обсудите с одноклассниками и с преподавателем следующие вопросы:</p><p><strong>1.</strong> Можно ли считать, что эксперимент группы Ленски действительно привёл весомые доказательства в пользу справедливости синтетической теории эволюции, или всё-таки результатов этого эксперимента недостаточно для того, чтобы считать эти положения доказанными? Приведите аргументы в пользу обеих точек зрения.</p><p><strong>2.</strong> Противники эволюционной теории достаточно часто говорят о том, что новый признак у организма не может появиться в результате мутации. Как вы думаете, можно ли опровергнуть данное утверждение, приведя в пример некоторые результаты эксперимента группы Ленски?</p>"
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            "text" => "<p><strong>1.</strong></p><div class="table-wrap"><table style="border: 1px solid rgb(0, 0, 0);"><tbody><tr><th style="border: 1px solid rgb(0, 0, 0);">Аргументы в пользу того, что эксперимент является весомым доказательством</th><th style="border: 1px solid rgb(0, 0, 0);">Аргументы в пользу того, что результатов эксперимента недостаточно</th></tr><tr><td style="border: 1px solid rgb(0, 0, 0);">Эксперимент является прямым наблюдением эволюции в реальном времени. Он наглядно демонстрирует действие таких факторов, как случайные мутации, естественный отбор и генетический дрейф, приводящих к адаптации популяции к условиям среды.</td><td style="border: 1px solid rgb(0, 0, 0);">Эксперимент проводился на одном виде асексуальных бактерий (<em>E. coli</em>) в крайне упрощённой и стабильной лабораторной среде. Результаты сложно экстраполировать на сложные, многоклеточные, половым путем размножающиеся организмы в изменчивых природных экосистемах.</td></tr><tr><td style="border: 1px solid rgb(0, 0, 0);">В ходе эксперимента зафиксировано появление нового, сложного признака — способности утилизировать цитрат в аэробных условиях. Это доказывает, что мутации могут приводить к появлению принципиально новых полезных функций, а не только к потере существующих.</td><td style="border: 1px solid rgb(0, 0, 0);">Эксперимент демонстрирует микроэволюцию (изменения внутри вида), но не макроэволюцию (возникновение новых видов, родов и т.д.). За десятки тысяч поколений не возникло ни одного нового вида бактерий, что, по мнению критиков, ограничивает значение эксперимента.</td></tr><tr><td style="border: 1px solid rgb(0, 0, 0);">Результаты показали, что эволюция не всегда предсказуема. Появление способности усваивать цитрат произошло только в одной из 12 популяций, что указывает на роль случайности и исторической обусловленности (контингентности) в эволюционном процессе.</td><td style="border: 1px solid rgb(0, 0, 0);">Несмотря на появление нового признака, он возник на основе уже существующих генов (через дупликацию и изменение регуляции гена-транспортера цитрата). Критики утверждают, что это не пример возникновения принципиально новой генетической информации, а лишь модификация старой.</td></tr></tbody></table></div><p><strong>2.</strong> Данное утверждение можно опровергнуть, приведя в пример ключевой результат эксперимента группы Ленски — появление у одной из популяций бактерий <em>E. coli</em> способности питаться цитратом в присутствии кислорода. Исходно бактерии этого вида не могут использовать цитрат в качестве источника энергии в аэробных условиях, и этот признак является одним из видовых определителей. Однако примерно через 31 500 поколений в одной из 12 пробирок бактерии приобрели эту новую способность. Генетический анализ показал, что это произошло в результате серии мутаций. Сначала произошла мутация-предпосылка (потенцирующая мутация), которая сама по себе не давала нового признака, но создавала условия для его появления. Затем произошла дупликация гена, кодирующего белок-транспортер цитрата, и эта копия попала под контроль нового промотора, что позволило гену работать в присутствии кислорода. Таким образом, эксперимент напрямую продемонстрировал, что случайные мутации могут приводить к возникновению совершенно нового и полезного для организма признака, который отсутствовал у предковых форм.</p>"
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            "text" => "<p><strong>1.</strong></p><div class="table-wrap"><table style="border: 1px solid rgb(0, 0, 0);"><tbody><tr><th style="border: 1px solid rgb(0, 0, 0);">Аргументы в пользу того, что эксперимент является весомым доказательством</th><th style="border: 1px solid rgb(0, 0, 0);">Аргументы в пользу того, что результатов эксперимента недостаточно</th></tr><tr><td style="border: 1px solid rgb(0, 0, 0);">Эксперимент является прямым наблюдением эволюции в реальном времени. Он наглядно демонстрирует действие таких факторов, как случайные мутации, естественный отбор и генетический дрейф, приводящих к адаптации популяции к условиям среды.</td><td style="border: 1px solid rgb(0, 0, 0);">Эксперимент проводился на одном виде асексуальных бактерий (<em>E. coli</em>) в крайне упрощённой и стабильной лабораторной среде. Результаты сложно экстраполировать на сложные, многоклеточные, половым путем размножающиеся организмы в изменчивых природных экосистемах.</td></tr><tr><td style="border: 1px solid rgb(0, 0, 0);">В ходе эксперимента зафиксировано появление нового, сложного признака — способности утилизировать цитрат в аэробных условиях. Это доказывает, что мутации могут приводить к появлению принципиально новых полезных функций, а не только к потере существующих.</td><td style="border: 1px solid rgb(0, 0, 0);">Эксперимент демонстрирует микроэволюцию (изменения внутри вида), но не макроэволюцию (возникновение новых видов, родов и т.д.). За десятки тысяч поколений не возникло ни одного нового вида бактерий, что, по мнению критиков, ограничивает значение эксперимента.</td></tr><tr><td style="border: 1px solid rgb(0, 0, 0);">Результаты показали, что эволюция не всегда предсказуема. Появление способности усваивать цитрат произошло только в одной из 12 популяций, что указывает на роль случайности и исторической обусловленности (контингентности) в эволюционном процессе.</td><td style="border: 1px solid rgb(0, 0, 0);">Несмотря на появление нового признака, он возник на основе уже существующих генов (через дупликацию и изменение регуляции гена-транспортера цитрата). Критики утверждают, что это не пример возникновения принципиально новой генетической информации, а лишь модификация старой.</td></tr></tbody></table></div><p><strong>2.</strong> Данное утверждение можно опровергнуть, приведя в пример ключевой результат эксперимента группы Ленски — появление у одной из популяций бактерий <em>E. coli</em> способности питаться цитратом в присутствии кислорода. Исходно бактерии этого вида не могут использовать цитрат в качестве источника энергии в аэробных условиях, и этот признак является одним из видовых определителей. Однако примерно через 31 500 поколений в одной из 12 пробирок бактерии приобрели эту новую способность. Генетический анализ показал, что это произошло в результате серии мутаций. Сначала произошла мутация-предпосылка (потенцирующая мутация), которая сама по себе не давала нового признака, но создавала условия для его появления. Затем произошла дупликация гена, кодирующего белок-транспортер цитрата, и эта копия попала под контроль нового промотора, что позволило гену работать в присутствии кислорода. Таким образом, эксперимент напрямую продемонстрировал, что случайные мутации могут приводить к возникновению совершенно нового и полезного для организма признака, который отсутствовал у предковых форм.</p>"
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Обсуждаем (с. 23)
Условие. Обсуждаем (с. 23)
скриншот условия
Биология, 11 класс Учебник, авторы: Пасечник Владимир Васильевич, Каменский Андрей Александрович, Рубцов Александр Михайлович, Швецов Глеб Геннадьевич, Абовян Леван Арташесович, Гапонюк Зоя Георгиевна, издательство Просвещение, Москва, 2019, страница 23, Условие
Биология, 11 класс Учебник, авторы: Пасечник Владимир Васильевич, Каменский Андрей Александрович, Рубцов Александр Михайлович, Швецов Глеб Геннадьевич, Абовян Леван Арташесович, Гапонюк Зоя Георгиевна, издательство Просвещение, Москва, 2019, страница 23, Условие (продолжение 2)

Прочитайте статью об эксперименте группы Ленски в рубрике «Это интересно», после чего обсудите с одноклассниками и с преподавателем следующие вопросы:

1. Можно ли считать, что эксперимент группы Ленски действительно привёл весомые доказательства в пользу справедливости синтетической теории эволюции, или всё-таки результатов этого эксперимента недостаточно для того, чтобы считать эти положения доказанными? Приведите аргументы в пользу обеих точек зрения.

2. Противники эволюционной теории достаточно часто говорят о том, что новый признак у организма не может появиться в результате мутации. Как вы думаете, можно ли опровергнуть данное утверждение, приведя в пример некоторые результаты эксперимента группы Ленски?

Решение 1. Обсуждаем (с. 23)
Биология, 11 класс Учебник, авторы: Пасечник Владимир Васильевич, Каменский Андрей Александрович, Рубцов Александр Михайлович, Швецов Глеб Геннадьевич, Абовян Леван Арташесович, Гапонюк Зоя Георгиевна, издательство Просвещение, Москва, 2019, страница 23, Решение 1
Биология, 11 класс Учебник, авторы: Пасечник Владимир Васильевич, Каменский Андрей Александрович, Рубцов Александр Михайлович, Швецов Глеб Геннадьевич, Абовян Леван Арташесович, Гапонюк Зоя Георгиевна, издательство Просвещение, Москва, 2019, страница 23, Решение 1 (продолжение 2)
Решение 2. Обсуждаем (с. 23)

1.

Аргументы в пользу того, что эксперимент является весомым доказательствомАргументы в пользу того, что результатов эксперимента недостаточно
Эксперимент является прямым наблюдением эволюции в реальном времени. Он наглядно демонстрирует действие таких факторов, как случайные мутации, естественный отбор и генетический дрейф, приводящих к адаптации популяции к условиям среды.Эксперимент проводился на одном виде асексуальных бактерий (E. coli) в крайне упрощённой и стабильной лабораторной среде. Результаты сложно экстраполировать на сложные, многоклеточные, половым путем размножающиеся организмы в изменчивых природных экосистемах.
В ходе эксперимента зафиксировано появление нового, сложного признака — способности утилизировать цитрат в аэробных условиях. Это доказывает, что мутации могут приводить к появлению принципиально новых полезных функций, а не только к потере существующих.Эксперимент демонстрирует микроэволюцию (изменения внутри вида), но не макроэволюцию (возникновение новых видов, родов и т.д.). За десятки тысяч поколений не возникло ни одного нового вида бактерий, что, по мнению критиков, ограничивает значение эксперимента.
Результаты показали, что эволюция не всегда предсказуема. Появление способности усваивать цитрат произошло только в одной из 12 популяций, что указывает на роль случайности и исторической обусловленности (контингентности) в эволюционном процессе.Несмотря на появление нового признака, он возник на основе уже существующих генов (через дупликацию и изменение регуляции гена-транспортера цитрата). Критики утверждают, что это не пример возникновения принципиально новой генетической информации, а лишь модификация старой.

2. Данное утверждение можно опровергнуть, приведя в пример ключевой результат эксперимента группы Ленски — появление у одной из популяций бактерий E. coli способности питаться цитратом в присутствии кислорода. Исходно бактерии этого вида не могут использовать цитрат в качестве источника энергии в аэробных условиях, и этот признак является одним из видовых определителей. Однако примерно через 31 500 поколений в одной из 12 пробирок бактерии приобрели эту новую способность. Генетический анализ показал, что это произошло в результате серии мутаций. Сначала произошла мутация-предпосылка (потенцирующая мутация), которая сама по себе не давала нового признака, но создавала условия для его появления. Затем произошла дупликация гена, кодирующего белок-транспортер цитрата, и эта копия попала под контроль нового промотора, что позволило гену работать в присутствии кислорода. Таким образом, эксперимент напрямую продемонстрировал, что случайные мутации могут приводить к возникновению совершенно нового и полезного для организма признака, который отсутствовал у предковых форм.

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